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I very nearly went with Alex Kingston here, and considered Lalla Ward and Mary Tamm as well, but honestly, I think Catherine Tate is the winner.

I'm American, so had no idea who Catherine Tate was when she turned up at the end of "Doomsday"; it wasn't until fandom exploded with ZOMG NOT THE COMEDIAN that I realized she was a big deal. But I loved her in TRB and was excited to see her in S4 – in addition to her terrific comedy sensibilities, she brought a lot of subtlety and emotion to a character who could have just been comic relief in the hands of someone less skilled. And while Donna may have gotten a shitty ending, Tate's performance that series ensured I'll give anything she's in a try.
Day 01 - Your Favorite Quote
Day 02 – Your Favorite Classic Series Episode
Day 03 – Your Favorite New Series Episode
Day 04 – Your Favorite Doctor
Day 05 – Your Favorite Companion
Day 06 – Whatever Tickles Your Fancy
Day 07 – Your Favorite Piece of Music
Day 08 – A Who-Related Photo That Makes You Happy
Day 09 – A Who-Related Photo That Makes You Angry/Sad
Day 10 – A Who-Related Photo That You Took
Day 11 – Your Favorite Season (Classic or New)
Day 12 – Whatever Tickles Your Fancy
Day 13 – Your Favorite Villain
Day 14 – The Villain Who Scared You the Most
Day 15 – Favorite Who-Related Tumblr
Day 16 – Your Favorite Who-Related FanFic
Day 17 – A Piece of Who-Related FanArt
Day 18 – Whatever Tickles Your Fancy
Day 20 – The Character Who Is Most Like You
Day 21 – Your Doctor Who OTP
Day 22 – A Who-Related Fan-Site
Day 23 – A Who-Related YouTube Video
Day 24 – Whatever Tickles Your Fancy
Day 25 – Favorite Who Actor
Day 26 – Favorite Who Actress
Day 27 – An Episode You Wish Hadn’t Been Made
Day 28 – An Episode Idea You Created Yourself
Day 29 – Who You Think Should Be the Next Doctor
Day 30 – Whatever Tickles Your Fancy
I'm American, so had no idea who Catherine Tate was when she turned up at the end of "Doomsday"; it wasn't until fandom exploded with ZOMG NOT THE COMEDIAN that I realized she was a big deal. But I loved her in TRB and was excited to see her in S4 – in addition to her terrific comedy sensibilities, she brought a lot of subtlety and emotion to a character who could have just been comic relief in the hands of someone less skilled. And while Donna may have gotten a shitty ending, Tate's performance that series ensured I'll give anything she's in a try.
Day 02 – Your Favorite Classic Series Episode
Day 03 – Your Favorite New Series Episode
Day 04 – Your Favorite Doctor
Day 05 – Your Favorite Companion
Day 06 – Whatever Tickles Your Fancy
Day 07 – Your Favorite Piece of Music
Day 08 – A Who-Related Photo That Makes You Happy
Day 09 – A Who-Related Photo That Makes You Angry/Sad
Day 10 – A Who-Related Photo That You Took
Day 11 – Your Favorite Season (Classic or New)
Day 12 – Whatever Tickles Your Fancy
Day 13 – Your Favorite Villain
Day 14 – The Villain Who Scared You the Most
Day 15 – Favorite Who-Related Tumblr
Day 16 – Your Favorite Who-Related FanFic
Day 17 – A Piece of Who-Related FanArt
Day 18 – Whatever Tickles Your Fancy
Day 20 – The Character Who Is Most Like You
Day 21 – Your Doctor Who OTP
Day 22 – A Who-Related Fan-Site
Day 23 – A Who-Related YouTube Video
Day 24 – Whatever Tickles Your Fancy
Day 25 – Favorite Who Actor
Day 26 – Favorite Who Actress
Day 27 – An Episode You Wish Hadn’t Been Made
Day 28 – An Episode Idea You Created Yourself
Day 29 – Who You Think Should Be the Next Doctor
Day 30 – Whatever Tickles Your Fancy